The Origin and Evolution of Life on Earth
Why do we start the discussion of biology with a discussion of geology?
Here are some reasons why one would do this:
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All life on earth is linked to geology; likewise, many surface geologic
features and processes have been influenced by life on earth. The origin
and early evolution of the earth are especially important when looking
at the origin and early evolution of life on earth.
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Life chemistry had its origin with the elements available to it during
the formation of the earth. These provided the basic raw materials available
for life.
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The evolution of life is in response to changes in environment. Many of
these changes are linked to changes in climate and/or geology.
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All life is (so far) limited to placement upon the earth, so it is crucial
that we understand the processes of the earth.
The Big Bang theory of the formation of the universe
All material in the universe was created in a huge "explosion," creating
and defining matter and space. The sudden cooling of the superheated ejecta
facilitated the combination of atomic components into atoms and molecules.
These clouds of gasses eventually cooled and formed the principle components
of galaxies - including stars and planets.
Other theories have been postulated (i.e. the oscillating universe theory
which states that the universe expands and contracts in a cycle every 100
billion years) and these are hotly debated
Location of the Solar System
Our solar system is located on a spiral arm of the Milky Way galaxy. We
are approximately 2/3 of the distance from the core to the outer rim of
the galaxy. This distance places us in a lower-density region of the galaxy
- there are some stars near to ours, but this region is nowhere as congested
as the galactic core.
Formation of the solar system
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A. The earth formed approximately 4.6 BYA (billion years ago.) Initially,
there was a cloud of gasses and dust particles, possibly originating from
the ejected particles of a nearby supernova.
B. The cloud gradually contracted and flattened, concentrating about
99% of its mass in the center with the rest rotating counterclockwise in
a flattened disk.
C. As the disk rotated, turbulence was created, causing condensation
of the disk into small, turbular eddies. These gradually accreted together
to form protoplanets.
D. These protoplanets further accreted, creating the mature planets
of the solar system.
The sun also accreted, pulling in most of the mass. As these accumulated,
the pressure and temperature caused the initiation of thermonuclear fusion.
This thermonuclear fusion is what provides the ultimate source of energy
for all life on earth.
In the hot accretions of planetesimals, iron-rich elements condensed
first, creating the cores (the inner and outer cores). Next, the lower-density
silicates began to condense and aggregate, forming the mantle and the crust.
Further differentiation of the crust was fueled by the energy output from
radioactive decay deep within the earth |
Structure of the earth
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There are three clearly defined regions of the earth:
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Cores (inner (1250 km) and outer (2100 km)) - composed primarily of iron
(85%) and nickel. Other heavy elements (such as radioactive elements) are
also found here
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Mantle (2900 km) - a "fluid" region primarily composed of oxygen and silica
- derived minerals
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Crust (5-70 km) - thin film of "crud" which has floated to the surface.
We are simply passengers, living on this geologic flotsam.
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The Dynamic Earth: Plate Tectonics
The outermost shell of the earth (the lithosphere) is a mosaic of twenty
or so large, rigid slabs of rock called tectonic
plates
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These plates move relative to each other at speeds measured in centimeters
per year
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This is the approximately the rate at which our fingernails grow
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The plates move very slowly on top of the layer of hotter, softer mantle
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Mantle is a liquid - but not in the sense that you think of a liquid
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It is similar to glass
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Where the plates grind against each other, stress builds up and is relieved
intermittently through earthquakes
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The areas where plates move are called faults
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Near plate boundaries, molten magma can rise to the surface and erupt to
form volcanoes
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Plate Interactions - Divergent Plate Boundaries
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The process of adjacent plates moving away from each other is best seen
at the oceanic spreading ridges
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New crust spews out from extensive oceanic ridge systems, like the one
seen in the middle of the Atlantic Ocean
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Smaller scale systems can be seen in rift valleys, such as the African
Rift Valley
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Divergent plate boundaries are the only source of new oceanic floor
Plate Interactions - Convergent Plate Boundaries
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When two plates meet, a convergent plate boundary forms
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Usually one plate will slide underneath the other - this will cause some
mountain building and lots of volcanic activity
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The Andes mountains in Chile are a classic example of this
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Sometimes plates will collide but one will not "smoothly" flow under another
- extensive and rapid mountain building will occur
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The Himalayans are an example of this
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The Movement of the Plates
The plates which form the terrestrial land masses have drifted together
and apart many times in the history of life on earth

The effects of these movements on life on earth cannot be underestimated.
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Climate is tied to the position of the plates
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Ocean currents, heat flow, salinity, and oxygen leves are dependent upon
plate positions
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Glaciation is tied to plate position
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Migration is possible if the plates are close together and isolation is
possible if the plates are far apart
The origin of the atmosphere and hydrosphere
Volatile gasses were either accreted to the earth during its formation
or they were brought here by meteorites. These volatiles were released
during the many incidences of heating and melting of the crust. This process
is know as outgassing; most outgassing occurred within the first 1 billion
years of the earth's history. The primitive atmosphere was rich in CO2,
N2, with lesser amounts of CO, H2,
HCl, with traces of NH3 and CH4.
Note that there is no O2 present in the
early atmosphere. Any O2 outgassed would have reacted
with the metals of the crust, causing oxidation. This lack of O2
is crucial for the formation of organic molecules!
What is life?
What is life? Life is everywhere, but it resists a simple one-sentence
definition. Some of the properties of life are given below:
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Order Living organisms partition resources and nutrients within
their systems. This is an energy-requiring process which must be maintained
for life to continue.
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Reproduction Organisms reproduce their own kind. Life only comes
from life.
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Growth and Development Heritable characters direct the pattern of
growth and development, producing an organism that is characteristic of
its species.
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Energy Utilization Organisms take in energy and transform it to
do work. Almost all of life's functions require energy.
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Homeostasis Regulatory mechanisms maintain an organism's internal
environment within tolerable limits, even though the external environment
may fluctuate. This process is known as homeostasis.
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Evolutionary Adaptation Live evolves as a result of the interaction
between organisms and their environment. As the environment is rarely stable,
life must adapt to survive in these new living conditions.
The Origin of Life - The Miller-Urey Experiment
How did life on earth first evolve? This question has plagued scientists
and philosophers for eons. While we still do not know, experiments have
given us some insights to how this may have occurred.
Experiments concerning the origin of life must answer the following
questions:
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How were small organic molecules (amino acids, nucleic acids, lipids)
formed in the primitive earth environment?
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How were these small organic molecules joined together to form polymers
(long chains of organic molecules)?
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How were abiotically produced molecules segregated into droplets called
protobionts
that have chemical compositions different from their surroundings
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What was the origin of heredity?
In 1953, Stanley Miller and Harold Urey used an apparatus (basically a
still) to recreate what is proposed to be the primitive environment of
the earth. A warmed flask of water simulated the primitive oceans. The
atmosphere in the Miller-Urey model was composed of H20,
H2, CH4 (methane), and NH3
(ammonia). Sparks were discharged in the synthetic atmosphere to mimic
lightning. A condenser cooled the atmosphere, raining water and any dissolved
compounds back into the miniature sea.
The Miller-Urey Experimental Apparatus
As materials circulated through the apparatus, the solution in the flask
changed from clear to murky brown. After one week, Miller and Urey analyzed
the contents of the solution and found a variety of organic compounds,
including amino acids, nucleic acids, and ATP, providing one scenario in
which the first criterion could have been met.
The primitive atmosphere was reducing (electron adding) which
facilitated the joining of monomers into polymers. The addition of sand
and clays into apparatus similar to the Miller-Urey can generate longer
polymers. The clays and sand act as catalysts which facilitate the formation
of polymers.
Experiments show protobionts form spontaneously from abiotically formed
organic molecules. These protobionts have membranes which are selectively
permeable, store energy (voltage difference), etc.
Lastly, simple, self-replicating systems have been created in the lab.
These consist of RNA systems. RNA systems can be self-catalyzing and are
relatively unstable. This would lead to a rapid evolution.
Much in not known about the origins of life, nor will it probably ever
be known. It is a very passionately debated subject. However, this is what
makes it interesting and will assure the study of this science for many
years to come.
Links to cool related sites
To learn more about the earth or any other planet, visit the nine
planets homepage
For a more complete look at plate tectonics, check out The
Dynamic Earth, the Story of Plate Tectonics
Extraterrestrial life? Check out the SETI
(Search for Extra Terrestrial Intelligence) project or CNN's mars
life page
If you have any questions or comments, please feel free to e-mail
me.