ANDRES CAMPOS, Ph.D.

Postdoctoral Fellow

RESEARCH SUMMARY:

The flagellar system has been extensively studied in enteric bacteria (Salmonella typhimurium and Escherichia coli). However, some questions about the regulation of the genomic expresion and the relation of this system with others into the cell, still reminds unanswered. The flagellar regulon consists of 40 to 50 genes arranged in different operons that are transcriptionally ordered in a hierarchy. The first level operon of this hierarchy (master operon) is composed by flhD and flhC genes. Both FlhD and FlhC, the protein products of the master operon, are required for the expression of all other flagellar genes. A heterotetrameric complex formed by FlhD and FlhC acts as the flagellar specific trascriptional activator of the flagellar genes of the second level. There are lines of evidence that suggest that FlhD (but not FlhC) is involved in other systems: A relation between the transcriptional regulation of the flagellar genes, the acid stress response, and the cell division has been established. Given the fact that E. coli flhD mutant cells display an unusual pattern in cell division, it has been suggested that FlhD affects this process in this bacteria. Recently, the FlhD genetic target was identified as cadA, but the mechanism by FlhD affects the expression of cadA is still unknown. We are studying the role of FlhD in the expression of cadA. It is possible that flhD affect the expresion of this gene by itself or as a complex with another factor that remains unknow.

The study of the specificity binding of FlhD with the proteins which it interacts and how the specificity binding of these complexes are altered are subjects of my work. We are carrying out studies in order to discern the structure of FlhD as well the structure of the different complexes formed by this protein: the heterotetramer formed by FlhD/FlhC; the interaction of this complex with the DNA; and if it is the case, the complexes formed by FlhD that intervene in the alteration of the expression of the cdaA.

Molecular biology, genetic, and biochemistry approaches will be done in to carry out this line of research.


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